About 130 years ago, Hermann Helmholtz devoted a whole chapter (60 pages of fine print) of his book on musical acoustics, "On the Sensations of Tone" to temperaments. He doesn't even mention a harmonica.
.............I mean, I like explanations as well as the next guy, but some of this stuff is way deep and (frankly) over my head.
I suspect that it is lack of interest more than lack of ability to understand.
I once quizzed a failing high school student about a motorcycle that quickly passed through an intersection two blocks away. He knew the make, model, engine displacement, drive train details, etc. because motorcycles interested him and his schoolwork did not.
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